Thenaria (Endomyaria) with usually well developed, but sometimes small base. Column with more or less distinct verrucae in its upper part. No sphincter or a weak endodermal one. Tentacles of two sorts, marginal and discal. Marginal tentacles arranged in cycles, discal tentacles in radial series, the former are smooth or have lateral protuberances, the latter are very short, usually papilliform, rarely they are indistinct or even absent. Many perfect pairs of mesenteries. Retractors of mesenteries strong.
Definitions and contained taxa are those of Oscar Carlgren as published in 1949 : Kungl Svenska Vetenskapsakadamiens Handlingar, Series 4, Volume 1, Number 1